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Plants

[ Plant Photo Key]
[ BFS Plant List in Excel Format]
[Unconfirmed plant reports]

NOTES:

Status of the list. Please note that this is a “living” list, constantly being refined and updated. Plant species and generic circumscriptions are in constant flux as a result of molecular research, and we attempt to keep the list up-to-date via the Jepson Online Interchange for California Floristics. In addition, some plant groups at the BFS are particularly challenging and open to revision.

We have recently updated and corrected the plant list, and reported species whose presence has not been confirmed have been moved to a separate list. If you can confirm that any of these species occur (or have occurred) at the BFS, please notify the webmaster.

Conservation status. Only three plant species at the BFS have special conservation status. Berberis nevinii is on both the state and federal Endangered Species lists (State and Federally Listed Endangered, Threatened, and Rare Plants of California, July 2016, State of California Department of Fish & Wildlife). Horkelia cuneata ssp. puberula is listed as “seriously endangered in California” by the California Native Plant Society (CNPS), although this designation carries no legal protection. CNPS also notes that Lepidium virginicum var. robinsonii is “uncommon in California”, but is not considered very endangered.

Origin. The origin listing indicates whether a plant is native to the site, a California native that has naturalized or been planted on the BFS, or exotic (not native to California).

Source. In general, we have credited plant identification to more recent sightings, although others, in addition to those cited, may have also identified the presence of a given species.

Photos & descriptions. ‘BFS’ links will show plants photographed at the BFS. We eventually hope to have photographs of all the plants on the site. Photos on the BFS pages may be used for any personal or academic purposes as long as notification of use is sent to the BFS Webmaster and the image is clearly credited with ©[Name of photographer]. For higher-resolution images, please contact the photographer. Note that no photos taken at the BFS may be used for commercial purposes without the express permission of the Claremont University Consortium.

‘CP’ links will return all photographs of the listed taxon in the CalPhotos, Digital Library Project database. As indicated above, some of these taxa may not, in fact, occur on the BFS. Note that you must check the stipulations with each photograph for permissible uses of the CalPhotos images. ‘Jep’ links will return the treatment and bioregional distribution for a taxon from Jepson eFlora, which parallels The Jepson Manual: Vascular Plants of California, 2nd Edition.

Herbarium specimens. Voucher specimens for a number of BFS plants have been deposited in several California herbaria, and the links will show the online data for those specimens.

Exotic plants around the old infirmary. The area surrounding the former Claremont Colleges’ infirmary building was once planted with a wide variety of plants, most of which are exotic. We have not attempted to identify all the exotic plants in this area, but the ones we have identified are included on the list.


Common name Species name (and notes) Origin* Source* Photos &
Descriptions*
Herbarium specimens*

DICOTS

Muskroot Family (Adoxaceae)

Blue Elderberry Sambucus nigra ssp. caerulea

Formerly Sambucus mexicana. Formerly placed in Caprifoliaceae.

native RSA, JG BFS, CP, Jep RSA549568
RSA611612
UCR107790
UC1595289

(all as Sambucus mexicana)

Amaranth Family (Amaranthaceae)

Tumble Pigweed, Tumbleweed Amaranthus albus exotic S, SD BFS, CP, Jep
Procumbent Pigweed Amaranthus blitoides native JW CP, Jep RSA766851
UCR224097

Sumac or Cashew Family (Anacardiaceae)

Laurel Sumac Malosma laurina native JG, S BFS, CP, Jep
Skunk Bush Rhus aromatica

Formerly Rhus trilobata.

native RSA, JG BFS, CP, Jep RSA549571
RSA614155
RSA764411
UC1595388

(above all as Rhus trilobata)

UCR107775
Lemonade Berry Rhus integrifolia native JG, S BFS, CP, Jep RSA549577
RSA614156
UCR108200
Sugar Bush Rhus ovata native RSA, JG BFS, CP, Jep RSA549578
UC1595298
Peruvian Pepper Tree Schinus molle exotic JG, S BFS, CP, Jep RSA611984
Western Poison Oak Toxicodendron diversilobum native JG, S BFS, CP, Jep RSA614157

Carrot Family (Apiaceae)

Wild Celery Apiastrum angustifolium native JG, SD BFS, CP, Jep RSA748786
UCR108172
UCR207701
Rattlesnake Weed Daucus pusillus native JG, SD BFS, CP, Jep RSA616156

Dogbane Family (Apocynaceae )

Kotolo, Indian Milkweed Asclepias eriocarpa

Formerly placed in Asclepiadaceae.

native S, SD BFS, CP, Jep RSA764402
Climbing Milkweed Funastrum cynanchoides var. hartwegii

Formerly Sarcostemma cynanchoides ssp. hartwegii. Formerly placed in Asclepiadaceae.

native S BFS, CP, Jep
Common Oleander Nerium oleander

One on east side of lake and several along south fence.

exotic S, SD BFS, CP, Jep

Sunflower Family (Asteraceae)

Annual Bur-Sage Ambrosia acanthicarpa native JG, SD CP, Jep
California Sagebrush Artemisia californica native JG, S BFS, CP, Jep
Mugwort Artemisia douglasiana native SD BFS, CP, Jep
Coyote Brush Baccharis pilularis native S, SD BFS, CP, Jep RSA549569
Mule Fat Baccharis salicifolia native RSA, JG BFS, CP, Jep RSA611982
RSA612652
RSA799914
UCR108461
Common Beggar-Ticks Bidens pilosa exotic S BFS, CP, Jep RSA823057
California Brickellbush Brickellia californica native SD, S BFS, CP, Jep
Italian Thistle Carduus pycnocephalus exotic JG, SD BFS, CP, Jep RSA751114
Tocalote, Maltese Star-Thistle Centaurea melitensis exotic S, SD BFS, CP, Jep
White Pincushion Chaenactis artemisiifolia native S BFS, CP, Jep
California Thistle, Cobwebby Thistle Cirsium occidentale

Several varieties may be present

native S, SD BFS, CP, Jep
Bull Thistle Cirsium vulgare

A small patch east of the lake has been eliminated.

exotic S BFS, CP, Jep
California Aster Corethrogyne filaginifolia

Formerly Lessingia filaginifolia var. filaginifolia. The Jepson Manual no longer recognizes any infraspecific taxa.

native JG, SD BFS, CP, Jep RSA611975
Australian Cotula, Southern Brass Buttons Cotula australis exotic VS CP, Jep RSA616147
African Daisy, Namaqualand Daisy Dimorphotheca sinuata

Several plants were found in the area burned in the Foothill Fire.

exotic S BFS, CP, Jep
California Brittlebush Encelia californica native* SD BFS, CP, Jep
Brittlebush, Incienso Encelia farinosa native* SD BFS, CP, Jep
Pine-Bush Ericameria pinifolia native RSA, JG BFS, CP, Jep RSA764413
UCR108196
Horseweed Erigeron canadensis

Formerly Conyza canadensis.

native EN, M BFS, CP, Jep
Leafy Fleabane Erigeron foliosus var. foliosus native S BFS, CP, Jep
Golden-Yarrow Eriophyllum confertiflorum native RSA, JG BFS, CP, Jep RSA612655
RSA751117
UCR108288
Saw-toothed Goldenbush Hazardia squarrosa R, S BFS, CP, Jep
Common Sunflower Helianthus annuus native S BFS, CP, Jep
Bristly Ox-Tongue Helminthotheca echioides

A single individual that appeared in the area burned by the Foothill Fire was removed.

exotic S BFS, CP, Jep RSA817064
Telegraph Weed Heterotheca grandiflora native EN, S, SD BFS, CP, Jep RSA764412
Smooth Cat’s-Ear Hypochaeris glabra exotic VS, DC BFS, CP, Jep
Prickly Lettuce Lactuca serriola exotic S, DC BFS, CP, Jep
California Goldfields Lasthenia californica

Has appeared transiently as a waif; not always present.

native RSA, JG CP, Jep RSA616152
Hawkweed Layia Layia hieracioides

To date only one small population near the north shore of pHake Lake is known at the BFS. This represents a range extension for this species.

native SD BFS, CP, Jep
Scale-Broom Lepidospartum squamatum native JG, S BFS, CP, Jep RSA611981
Valley Lessingia Lessingia glandulifera var. glandulifera native VS, SD BFS, CP, Jep RSA611968
Daggerleaf Cottonrose Logfia gallica

Formerly Filago gallica.

exotic VS CP, Jep RSA616158

(as Filago gallica)

Cudweed, Two-tone Everlasting Pseudognaphalium biolettii

Formerly Gnaphalium bicolor.

native RSA, JG BFS, CP, Jep UCR108175

(as Gnaphalium bicolor)

California Cudweed Pseudognaphalium californicum

Formerly Gnaphalium californicum

native EN, JG BFS, CP, Jep RSA751109

(as Gnaphalium californicum)

Woolly-Heads, Slender Woolly-Marbles Psilocarphus tenellus native EN, R CP, Jep
Threadleaf Ragwort Senecio flaccidus var. douglasii native S BFS, CP, Jep RSA444578
Common Groundsel Senecio vulgaris exotic S BFS, CP, Jep RSA794414
Milk Thistle Silybum marianum

In 2010 a large clump of Milk Thistle was removed from the in parkway on Mills Ave. It has not since recurred.

exotic S, SD BFS, CP, Jep
Prickly Sow Thistle Sonchus asper exotic S, SD BFS, CP, Jep
Common Sow Thistle Sonchus oleraceus exotic LG CP, Jep
Wand Wirelettuce Stephanomeria virgata ssp. pleurocarpa native EN, JG, M BFS, CP, Jep RSA611972
Rod Wirelettuce Stephanomeria virgata ssp. virgata native S BFS, CP, Jep RSA823055
Everlasting Nest Straw Stylocline gnaphaloides native VS BFS, CP, Jep RSA612073
Cotton-Thorn Tetradymia comosa native JG, S BFS, CP, Jep RSA549566
RSA611974
RSA764414
UC1595291

Barberry Family (Berberidaceae)

Nevin’s Barberry Berberis nevinii native S, SD BFS, CP, Jep RSA792777

Birch or Alder Family (Betulaceae)

White Alder Alnus rhombifolia native* JG, S BFS, CP, Jep RSA611613
RSA766852

Borage Family (Boraginaceae)

Common Fiddleneck Amsinckia intermedia

Formerly Amsinckia menziesii var. intermedia

native SD BFS, CP, Jep POM612662

(as Amsinckia menziesii var. intermedia)

UCR108167
Bearded Cryptantha Cryptantha barbigera native KH CP, Jep
Popcorn Flower Cryptantha intermedia native SD BFS, CP, Jep RSA612076
RSA612663
RSA751118
Redroot Cryptantha Cryptantha micrantha native KH CP, Jep
Minute-flowered Cryptantha Cryptantha micromeres native KH CP, Jep
Common Cryptantha Cryptantha muricata native RSA, JG CP, Jep
Prickly-Nut Cryptantha Cryptantha muricata var. denticulata native KH CP, Jep
Rigid Cryptantha Cryptantha nevadensis var. rigida native KH CP, Jep
Whispering Bells Emmenanthe penduliflora var. penduliflora native S BFS, CP, Jep
Hairy Yerba Santa Eriodictyon trichocalyx var. trichocalyx

Formerly placed in Hydrophyllaceae.

native RSA, JG BFS, CP, Jep RSA611616
UCR108201
Common Eucrypta Eucrypta chrysanthemifolia var. chrysanthemifolia

Formerly placed in Hydrophyllaceae.

native RSA, JG BFS, CP, Jep RSA611616
RSA549583
RSA612172
Alkali Heliotrope, Seaside Heliotrope Heliotropium curassavicum var. oculatum

Formerly Heliotropium curassavicum.

native† SD BFS, CP, Jep
Comb-Bur, Narrow-toothed Pectocarya Pectocarya linearis ssp. ferocula native JG, SD CP, Jep RSA616160B
RSA792784
Sleeping Combseed, Northern Pectocarya Pectocarya penicillata native VS CP, Jep RSA612072
RSA616160A
Desert Bluebells Phacelia campanularia

In Mills Ave parkway. Not native to the BFS; probably planted in a wildflower mix.

native* S BFS, CP, Jep
Caterpillar Phacelia Phacelia cicutaria var. hispida

Formerly placed in Hydrophyllaceae.

native S BFS, CP, Jep
Distant Phacelia Phacelia distans

Formerly placed in Hydrophyllaceae.

native RSA, JG BFS, CP, Jep RSA611615
RSA799926
UCR107754
California Bell, Wild Canterbury Bells Phacelia minor

Formerly placed in Hydrophyllaceae.

native RSA, JG BFS, CP, Jep RSA612070
RSA612173
RSA799923
Branching Phacelia Phacelia ramosissima

Formerly in placed Hydrophyllaceae.

native SD BFS, Jep RSA751108
Lacy Scorpionweed, Tansy Phacelia Phacelia tanacetifolia

Formerly placed in Hydrophyllaceae.

native EN BFS, CP, Jep
Popcorn Flower Plagiobothrys canescens native SD CP, Jep
Rough-stemmed Popcorn Flower Plagiobothrys collinus var. fulvescens native VS, SD CP, Jep RSA616155
RSA792783

Mustard Family (Brassicaceae)

Common Sandweed Athysanus pusillus native TR, JW CP, Jep RSA792772
UC1595297
Indian Mustard Brassica fruticulosa

Native to Europe, recently reported as a new introduction to North America, collected at nearby Evey Canyon and well established at the south foot of the San Gabriel and San Bernadino Mountains of Los Angeles, San Bernadino, and Riverside Counites. Keys to B. juncea in the treatment of the The Jepson Manual, differing in having more consistently lobed leaves, shorter beak (3-4 mm) of the fruit, strongly beaded appearance of the mature fruits, and smaller flowers. See Sanders, Madroño 43(4): 523-524, Oct-Dec 1996. A dense local infestation occurs in the northern portion of the field station.

exotic RSA, JG, JW Jep RSA792778
RSA831433
Black Mustard Brassica nigra exotic RSA, JG BFS, CP, Jep RSA611969
Turnip, Field Mustard Brassica rapa exotic RSA, JG BFS, CP, Jep
Sahara Mustard Brassica tournefortii

First detected in in the parkway along Foothill Blvd., particularly near the entry gate and along the frontage road between Dartmouth and College Aves. Despite eradication attempts, plants have been detected in the parkway, inside the entry gate, by the Abilene Way gate, and north of the infirmary.

exotic LG BFS, CP, Jep RSA781424
Bitter-cress, Toothwort Cardamine oligosperma native S, SD BFS, CP, Jep UC1595296
Shepherd’s Purse Capsella bursa-pastoris exotic SD BFS, CP, Jep
Western Tansy-Mustard Descurainia pinnata native JW BFS, CP, Jep RSA792779
Garden Rocket Eruca vesicaria ssp. sativa exotic S BFS, CP, Jep RSA831432
Short-pod Mustard Hirschfeldia incana exotic EN BFS, CP, Jep UCR108416
UCR107755

(both as Brassica geniculata)

Lesser Swine Cress Lepidium didymum

Formerly Coronopus didymus.

exotic S BFS, CP, Jep
Hairypod Pepperweed Lepidium lasiocarpum ssp. lasiocarpum native S CP, Jep RSA799917
Robinson’s Peppergrass Lepidium virginicum var. robinsonii

The Jepson Manual, 2nd ed., lumps L. virginicum var. robinsonii together with Lepidium virginicum var. pubesecens and Lepidium virginicum var. robinsonii into Lepidium virginicum var. menziesii.

native JS BFS, Jep
Sweet Alyssum Lobularia maritima exotic RSA, JG BFS, CP, Jep RSA549565
UCR108174
UC1595292
London Rocket Sisymbrium irio exotic SD CP, Jep
Indian Hedgemustard, Eastern Rocket Sisymbrium orientale exotic JW CP, Jep RSA748150
UCR207700

Cactus Family (Cactaceae)

Night-blooming Cereus, Peruvian Apple Cactus Cereus peruvianus

One planted outside the BFS Field House. A small group of plants naturalized from an adjacent yard on the eastern side of the ‘Neck’ has been removed.

exotic S, SD BFS
Cane or Snake Cholla Cylindropuntia californica

Formerly Opuntia parryi. The large genus Opuntia was subdivided into five genera; true chollas are in the genus Cylindropuntia, and O. parryi was renamed C. californica.

native RSA, JG BFS, CP, Jep RSA612661

(as Opuntia parryi)

Indian-Fig, Mission Prickly-Pear Opuntia ficus-indica exotic JG, S BFS, CP, Jep
Prickly-Pear Opuntia littoralis native JG, S BFS, CP, Jep RSA748071

Pink Family (Caryophyllaceae)

Spreading Pygmyleaf Loeflingia squarrosa

Formerly Loeflingia squarrosa var. squarrosa. The Jepson Manual no longer recognizes any infraspecific taxa.

native VS CP, Jep RSA612084
Four-leaved Allseed Polycarpon tetraphyllum exotic VS CP, Jep RSA616146
RSA616148
Sleepy Catchfly Silene antirrhina native S BFS, CP, Jep
Common Chickweed Stellaria media exotic EN CP, Jep

Goosefoot Family (Chenopodiaceae)

Four-wing Saltbush Atriplex canescens var. canescens

A single large specimen near the southwest corner of the ‘Neck’. Formerly in Amaranthaceae.

native† SD BFS, CP, Jep
Pitseed Goosefoot Chenopodium berlandieri

Formerly in Amaranthaceae.

native S BFS, CP, Jep UCD151063 RSA823056
California Goosefoot Chenopodium californicum

Formerly in Amaranthaceae.

native S, SD BFS, CP, Jep RSA799915
Russian Thistle Salsola australis exotic S BFS, CP, Jep RSA831430
Russian Thistle Salsola tragus exotic S BFS, CP, Jep RSA831431

Spiderflower Family (Cleomaceae)

Bladderpod Peritoma arborea

Formerly Cleome isomeris and Isomeris arborea. Formerly placed in Capparaceae and Brassicaceae.

native† S, SD BFS, CP, Jep RSA549570

(as Cleome isomeris)

RSA748781
UC1595389

(as Isomeris arborea)

Morning-Glory Family (Convolvulaceae)

Island False Bindweed, Morning-Glory Calystegia macrostegia subsp. intermedia native S, SD BFS, CP, Jep RSA831434
Chaparral Dodder Cuscuta californica

Formerly placed in Cuscutaceae.

native JG BFS, CP, Jep RSA751110

Stone Crop Family (Crassulaceae)

Pygmy-Weed Crassula connata native R CP, Jep
Moss Pygmyweed Crassula tillaea exotic VS CP, Jep RSA611986
RSA792834
Lance-leaved Dudleya Dudleya lanceolata native S BFS, CP, Jep

Cucumber Family (Cucurbitaceae)

Calabazilla, Buffalo Gourd Cucurbita foetidissima native VS, S BFS, CP, Jep
Man-Root, Wild Cucumber Marah macrocarpa

Formerly Marah macrocarpus var. macrocarpus. The Jepson Manual no longer recognizes any infraspecific taxa.

native RSA, JG BFS, CP, Jep RSA611611

Spurge Family (Euphorbiaceae)

California Croton Croton californicus native RSA, JG BFS, CP, Jep RSA611967
RSA612178
RSA751107
RSA764415
UCR108228
Dove Weed, Turkey Mullein Croton setiger

Formerly Eremocarpus setiger.

native JG, SD BFS, CP, Jep RSA611970
RSA764421

(both as Eremocarpus setigerus)

Rattlesnake Sandmat Euphorbia albomarginata

Formerly Chamaesyce albomarginata.

native R CP, Jep
Petty Spurge Euphorbia peplus exotic JG, SD BFS, CP, Jep RSA612078
Thyme-leafed Spurge Euphorbia serpyllifolia

Formerly Chamaesyce serpyllifolia.

native M BFS, CP, Jep
Castor Bean Ricinus communis exotic RSA, JG BFS, CP, Jep UCR107781

Legume (Bean & Pea) Family (Fabaceae)

Spanish Clover Acmispon americanus var. americanus

Formerly Lotus unifoliolatus var. unifoliolatus.

native S BFS, CP, Jep
Short-winged Deerweed Acmispon glaber var. brevialatus

Formerly Lotus scoparius var. brevialatus.

native RSA, JG CP, Jep UCR107830

(as Lotus scoparius var. brevialatus)

Deerweed Acmispon glaber var. glaber

Formerly Lotus scoparius var. scoparius.

native JG BFS, CP, Jep
San Diego Bird’s-foot-trefoil Acmispon micranthus

Formerly Lotus hamatus.

native VS CP, Jep RSA612083

(as Lotus hamatus)

Bishop Lotus Acmispon strigosus

Formerly Lotus strigosus.

native VS BFS, CP, Jep RSA612081
RSA616153

(as Lotus strigosus)

Bird-of-Paradise Shrub Caesalpinia gilliesii exotic DC BFS, CP, Jep
Western Redbud Cercis occidentalis native† SD BFS, CP, Jep
Miniature Lupine Lupinus bicolor native R, RSA, JG BFS, CP, Jep RSA612177
Sky Lupine Lupinus nanus

In Foothill Blvd parkway. Not native to the BFS; probably planted in a wildflower mix.

native* S BFS, CP, Jep
Coulter’s Lupine Lupinus sparsiflorus native S BFS, CP, Jep RSA817063
Arroyo Lupine Lupinus succulentus native S BFS, CP, Jep
Collar Lupine Lupinus truncatus native VS, S CP, Jep RSA612071
RSA799925
Sourclover Melilotus indicus exotic JG BFS, CP, Jep RSA615305
RSA615307
Mexican Palo Verde Parkinsonia aculeata

Naturalized in So. Cal.; one by front gate and one in wash near Lowell gate

exotic S, SD BFS, CP, Jep
Honey Mesquite Prosopis glandulosa native* JW BFS, CP, Jep RSA748784
Truncate Sack Clover Trifolium depauperatum var. truncatum native S BFS, CP, Jep RSA831435
Pinpoint Clover Trifolium gracilentum native VS CP, Jep RSA612065
Small-head Clover Trifolium microcephalum native VS CP, Jep RSA612082
RSA615303

Beech Family (Fagaceae)

Oak (unspecified) Quercus sp.

In experimental plots in East Field established by the Gamon lab

native* S CP, Jep RSA792774
RSA792775
Coast Live Oak Quercus agrifolia var. agrifolia native VS, S BFS, CP, Jep RSA612079
Scrub Oak Quercus berberidifolia native JG, JW BFS, CP, Jep RSA792836
Engelman Oak, Mesa Oak Quercus engelmannii native* SD CP, Jep
Valley Oak, Roble Quercus lobata

Two deciduous oaks planted east of the entrance road just south of the infirmary.

native* SD, S BFS, CP, Jep

Gentian Family (Gentianaceae)

Canchalagua, California Centaury, Charming Centaury Zeltnera venusta

Formerly Centaurium venustum

native JG, S BFS, CP, Jep UCR146987

(as Centaurium venustum)

Geranium Family (Geraniaceae)

Filaree, Storksbill Erodium botrys exotic VS CP, Jep RSA751116
Red-stem Filaree Erodium cicutarium exotic RSA, JG BFS, CP, Jep RSA612653
UCR107801
White-stem Filaree, Greenstem Filaree Erodium moschatum exotic S BFS, CP, Jep

Gooseberry or Currant Family (Grossulariaceae)

Bitter Gooseberry Ribes amarum native† TR CP, Jep RSA549572
Golden Currant Ribes aureum var. gracillimum native RSA, JG BFS, CP, Jep RSA748783
White-Flowering Currant Ribes indecorum native RSA, JG BFS, CP, Jep RSA549567
RSA612175
UCR107800
UC1595290
Fuchsia-flowered Gooseberry Ribes speciosum native† R BFS, CP, Jep RSA549573

Walnut Family (Juglandaceae)

Southern California Black Walnut Juglans californica

Formerly listed as Juglans californica var. californica, but the Jepson Manual no longer recognizes any infraspecific taxa.

native JG, S BFS, CP, Jep RSA764418
UCR224313

Mint Family (Lamiaceae)

Henbit, Giraffe Head, Dead Nettle Lamium amplexicaule exotic SD BFS, CP, Jep RSA792771
Horehound Marrubium vulgare exotic RSA, JG BFS, CP, Jep RSA613095
UCR108173
White Sage Salvia apiana native JG, S BFS, CP, Jep RSA613094
UCR107758
Chia Salvia columbariae native S BFS, CP, Jep
Black Sage Salvia mellifera native S, SD BFS, CP, Jep

Laurel Family (Lauraceae)

California Bay, California Laurel Umbellularia californica native* JG, SD BFS, CP, Jep RSA613093

Loosestrife Family (Lythraceae)

Hyssop Loosestrife Lythrum hyssopifolia exotic R, VS CP, Jep RSA616149

Mallow Family (Malvaceae)

Flannelbush Fremontodendron californicum

Formerly Fremontodendron californicum ssp. californicum; The Jepson Manual no longer recognizes any infraspecific taxa of this species. Formerly placed in Sterculiaceae.

native† JG, S BFS, CP, Jep UCR-108170
Chaparral Mallow Malacothamnus fasciculatus native SD BFS, CP, Jep

Mahogany Family (Meliaceae)

Chinaberry Melia azedarach exotic S BFS, CP, Jep

Miner’s Lettuce Family (Montiaceae)

Red Maids Calandrinia menziesii

Formerly Calandrinia ciliata. Formerly placed in Portulacaceae.

native S BFS, CP, Jep RSA799927

(as Calandrinia ciliata)

Streambank Springbeauty Claytonia parviflora ssp. parviflora

Formerly placed in Portulacaceae.

native S BFS, CP, Jep RSA748782

Myrsine Family (Myrsinaceae)

Scarlet Pimpernel Lysimachia arvensis

Formerly Anagallis arvensis. Formerly placed in Primulaceae.

exotic JG, S BFS, CP, Jep RSA616150

(as Anagallis arvensis)

Myrtle Family (Myrtaceae)

Blue Gum Eucalyptus globulus exotic S, SD BFS, CP, Jep
Red Iron Bark Eucalyptus sideroxylon exotic JW BFS, CP, Jep RSA748149

Four O’clock Family (Nyctaginaceae)

California Four O’Clock, Wishbone Bush Mirabilis laevis var. crassifolia

Formerly Mirabilis californica.

native RSA, JG BFS, CP, Jep UCR108169

(as Mirabilis californica)

RSA613090

Olive Family (Oleaceae)

Desert Olive Forestiera pubescens

A single male plant grows on the north shore of pHake Lake.

native† LG BFS, CP, Jep RSA778384
RSA781425
Single-Leaf Ash Fraxinus anomala native S BFS, CP, Jep
California Ash Fraxinus dipetala native EN CP, Jep
European Olive Olea europaea exotic EN, JG BFS, CP, Jep RSA748148

Evening Primrose Family (Onagraceae)

California Sun Cup Camissoniopsis bistorta

Formerly Camissonia bistorta

native RSA, JG BFS, CP, Jep RSA611985
RSA613089
RSA751111
RSA751112
RSA799913

(all as Camissonia bistorta)

Minature Sun Cup, Spencer Primrose Camissoniopsis micrantha

Formerly Camissonia micrantha

native S BFS, CP, Jep RSA817062
Four Spot Clarkia purpurea ssp. quadrivulnera native SD, S BFS, CP, Jep
Elegant Clarkia Clarkia unguiculata native† S BFS, CP, Jep
Willowherb Epilobium ciliatum native† S CP, Jep RSA766856
Water Primrose Ludwigia repens native SD BFS, Jep RSA764407

Peony Family (Paeoniaceae)

California peony Paeonia californica native SD, S BFS, CP, Jep RSA792776

Poppy Family (Papaveraceae)

California Poppy Eschscholzia californica native JG, S BFS, CP, Jep
Coulter’s Matilija Poppy Romneya coulteri
native*† SD, S BFS, CP, Jep RSA751106

Lopseed Family (Phrymaceae)

Sticky Monkey Flower Mimulus aurantiacus

Formerly placed in Scrophulariaceae.

native JG, S BFS, CP, Jep RSA612183

Plantain Family (Plantaginaceae)

Blue Toadflax Nuttallanthus texanus

Formerly Linaria canadensis. Formerly placed in Scrophulariaceae.

native VS, S BFS, CP, Jep RSA616159

(as Linaria canadensis)

Royal Penstemon Penstemon spectabilis

Formerly placed in Scrophulariaceae.

native RSA, JG BFS, CP, Jep
Royal Penstemon Penstemon spectabilis var. spectabilis

Formerly placed in Scrophulariaceae.

native RSA, JG CP, Jep RSA612181
UCR107832

Sycamore Family (Platanaceae)

Western Sycamore Platanus racemosa native RSA, JG BFS, CP, Jep RSA613088
UCR108191

Phlox Family (Polemoniaceae)

Chaparral Gilia Gilia angelensis native LG BFS, CP, Jep RSA781423
Sapphire Eriastrum Eriastrum sapphirinum native EN, JG BFS, CP, Jep RSA751113
Hollyleaf Navarretia Navarretia atractyloides native JW BFS, CP, Jep RSA751104

Buckwheat Family (Polygonaceae)

Leafy California Buckwheat Eriogonum fasciculatum var. foliolosum native JG, S BFS, CP, Jep RSA611976
RSA614149
UCR108197
Slender Wooly Wild Buckwheat Eriogonum gracile var. gracile native JG BFS, CP, Jep RSA611973
RSA612085
Leather Spineflower Lastarriaea coriacea native VS CP, Jep RSA612075
False Waterpepper Persicaria hydropiperoides

Formerly Polygonum hydropiperoides

native JW, LG CP, Jep RSA764408

(as Polygonum hydropiperoides)

RSA778386
Dotted Smartweed Persicaria punctata

Formerly Polygonum punctatum

native VS, LG BFS, CP, Jep RSA611980

(as Polygonum punctatum)

RSA778385
Annual Beard Grass, Rabbitfoot Grass Polygonum monspeliensis exotic VS Jep RSA616145
RSA764410
Woodland Threadstem Pterostegia drymarioides native JG, S BFS, CP, Jep RSA616157
Wild-Rhubarb, Canaigre Rumex hymenosepalus native RSA, JG BFS, CP, Jep RSA614148
UCR108199

Buttercup Family (Ranunculaceae)

Chaparral Clematis, Pipstem Clematis Clematis lasiantha native† JW BFS, CP, Jep RSA792835
Cardinal or Scarlet Larkspur Delphinium cardinale native EN, R, JG BFS, CP, Jep RSA612066
Parry’s Larkspur Delphinium parryi ssp. parryi native VS, M BFS, CP, Jep RSA612068

Coffeeberry Family (Rhamnaceae)

Hoaryleaf Ceanothus Ceanothus crassifolius

In experimental plots in East Field established by the Gamon lab

native* JW CP, Jep RSA792773
California Coffee Berry Frangula californica ssp. californica

Formerly Rhamnus californica ssp. californica.

native RSA, JG BFS, CP, Jep RSA614147
RSA764416
UCR107831

(all as Rhamnus californica)

Spiny Redberry Rhamnus crocea native RSA, JG BFS, CP, Jep RSA549576
UCR108198
UC1595300

Rose Family (Rosaceae)

Birch-leaf Mountain-Mahogany Cercocarpus betuloides native* R, S BFS, CP, Jep
Toyon, Christmas Berry Heteromeles arbutifolia native RSA, JG BFS, CP, Jep RSA611617
Mesa Horkelia, Wedge-Leaf Honeydew Horkelia cuneata var. puberula

Listed as seriously endangered in California by the California Native Plant Society. The poplulation at the BFS represents one of the few known in Los Angeles County.

native R, JG BFS, CP, Jep RSA611622
RSA612067
RSA748147
UCR109147
Holly-leafed Cherry, Islay Prunus ilicifolia ssp. ilicifolia

P. ilicifolia ssp. ilicifolia hybridizes freely with P. ilicifolia ssp. lyonii, and intergrading forms may exist at the BFS.

native RSA, JG BFS, CP, Jep RSA612184
RSA612185
UCR107833
Catalina Cherry Prunus ilicifolia ssp. lyonii

P. ilicifolia ssp. ilicifolia hybridizes freely with P. ilicifolia ssp. lyonii, and intergrading forms may exist at the BFS.

native* RSA, JG BFS, CP, Jep
Small-leaved Rose Rosa minutifolia native* SD CP, Jep

Madder Family (Rubiaceae)

Goose Grass Galium aparine native VS BFS, CP, Jep RSA612077
San Diego Bedstraw Galium nuttallii subsp. nuttallii native JW Jep RSA799917

Willow Family (Salicaceae)

Fremont Cottonwood, Alamo Cottonwood Populus fremontii ssp. fremontii native SD BFS, CP, Jep
Gooding’s Black Willow Salix gooddingii native EN, SD BFS, CP, Jep RSA766853
Arroyo Willow Salix lasiolepis native JG, SD BFS, CP, Jep RSA612651
RSA615308
RSA748780

Soapberry Family (Sapindaceae)

Chinese Flame-Tree Koelreuteria bipinnata exotic JW CP RSA764420

Figwort Family (Scrophulariaceae)

Mediterranean Figwort, Nettle-leaved Figwort Scrophularia peregrina

Previously unreported in North America, S. peregrina had become an established weed throughout the Rancho Santa Ana Botanic Garden by 1996 (Ross and Boyd, Madroño 43(4): 436, July-Sept 1996), and has since spread to the BFS. Native to Portugal and the Mediterranean region.

exotic RSA, JG BFS, CP, Jep RSA574289
RSA792782
UCR89484
UC1611552
Wand Mullein Verbascum virgatum exotic SD, S BFS, CP, Jep RSA751105

Simarouba Family (Simaroubaceae)

Tree of Heaven Ailanthus altissima

One infestation in the Foothill Blvd parkway was removed in 2011, but other scattered individuals may remain within the BFS.

exotic JW, S BFS, CP, Jep RSA764419

Nightshade Family (Solanaceae)

Jimson Weed Datura wrightii native JG, S BFS, CP, Jep
Manyflower Tobacco Nicotiana acuminata var. multiflora exotic SD, S BFS, CP, Jep
Tree Tobacco Nicotiana glauca exotic JG, S BFS, CP, Jep RSA611983
RSA613098
Indian Tobacco Nicotiana quadrivalvis native SD, S BFS, CP, Jep RSA748151
RSA764406
Douglas’ Nightshade Solanum douglasii native JG, SD BFS, CP, Jep RSA613099
RSA751115
RSA799916
Purple Nightshade, Bluewitch Solanum umbelliferum

Formerly listed as Solanum xanti, but S. xanti and S. umbelliferum have been consolidated as S. umbelliferum.

native TR, JG, SD BFS, CP, Jep UCR107761

(as Solanum xanti)

UC1595293
RSA549564
RSA613097

Nettle Family (Urticaceae)

Black-hair Nettle Hesperocnide tenella native† TR, JW CP, Jep RSA549581
RSA562724
RSA792781
UC1595295
Pellitory Parietaria hespera var. hespera native VS CP, Jep RSA615306
Dwarf Nettle Urtica urens exotic JG, S BFS, CP, Jep RSA549582
RSA792780
SEINET4118144
UC1595294

Mistletoe Family (Viscaceae)

Big Leaf Mistletoe Phoradendron leucarpum ssp. macrophyllum

Formerly Phoradendron macrophyllum and Phoradendron serotinum ssp. macrophyllum

native JG, SD BFS, CP, Jep

Grape Family (Vitaceae)

Wild Grape Vitis sp.

Because the few Vitis plants on the BFS have not borne fruit, species identification is difficult. Plants may be V. californica, V. girdiana, or a hybrid.

native SD, S BFS, CP, Jep

Caltrop Family (Zygophyllaceae)

Puncture Vine Tribulus terrestris

First detected in 2008 as an infestation in the northeast corner of the East Field. In spite of eradication attempts, plants have appeared in the East Field, near the entry gate, and in the Foothill Blvd Parkway.

native SD, S BFS, CP, Jep

MONOCOTS

Agave Family (Agavaceae)

American Agave, Century Plant Agave americana

Formerly placed in Liliaceae.

exotic S BFS, CP, Jep
Chaparral Yucca, Our Lord’s Candle Hesperoyucca whipplei

Formerly Yucca whipplei. Formerly placed in Liliaceae.

native S BFS, CP, Jep
California Soap Plant Chlorogalum pomeridianum var. pomeridianum

Formerly placed in Liliaceae.

native SD BFS, CP, Jep

Arum Family (Araceae)

Duckweed Lemna valdiviana

Formerly placed in Lemnaceae.

native VS BFS, CP, Jep RSA613092
Tongueshape Mud-Midget Wolffiella lingulata native LG Jep RSA613092
(in specimen of Lemna valdiviana)

Palm Family (Arecaceae)

California Fan Palm Washingtonia filifera native† SD, S CP, Jep RSA764404
Mexican Fan Palm Washingtonia robusta exotic S CP, Jep

Asparagus Family (Asparagaceae)

Bridal Creeper Asparagus asparagoides exotic LG BFS, CP, Jep

Sedge Family (Cyperaceae)

Tall Flatsedge Cyperus eragrostis native S CP, Jep RSA764405
Purple spikerush Eleocharis atropurpurea native SD CP, Jep
Spikerush Eleocharis macrostachya

Observed only in vernal pools to west of pHake Lake.

native VS CP, Jep RSA612180
Low Bulrush Isolepis cernua native JW, LG CP, Jep RSA766855
RSA778383
Naked-stemmed Bulrush Schoenoplectus sp.

Formerly listed as Scirpus sp., but California Scirpus are now placed in Amphiscirpus, Bolboschoenus, Isolepis, Schoenoplectus, or Trichophorum.

native SD CP, Jep

Iris Family (Iridaceae)

Western Blue-eyed Grass Sisyrinchium bellum

A small population near the southwest corner of pHake Lake.

native VS, SD BFS, CP, Jep RSA612174

Rush Family (Juncaceae)

Baltic Rush Juncus balticus var. ater

Clumped perennial with twisted stems and congested inflorescences afound on the margin of pHake Lake are tentatively identified as this species. [EN]

native EN CP, Jep
Toad Rush Juncus bufonius native JG CP, Jep
Sonoran Rush,
Baja Rush
Juncus effusus ssp. austrocalifornicus native VS Jep RSA611979
RSA764403
Mexican Rush Juncus mexicanus

Diffuse spreading plants with twisted stems, small open inflorescences, and leaves resembling stems are referred to this species, found only on the north shore of pHake Lake, near the roadway. [EN]

native JG CP, Jep RSA748779
Wrinkled Rush Juncus rugulosus native VS CP, Jep RSA611978
RSA764409

Grass Family (Poaceae)

Slender Wild Oat Avena barbata exotic R, JG CP, Jep RSA614152
UCR108415
SEINET3919640
Wild Oat Avena fatua exotic M CP, Jep
Ripgut Grass Bromus diandrus exotic EN, JG BFS, CP, Jep RSA614150
UCR108414
Soft Chess, Soft Brome Bromus hordeaceus exotic EN, R, JG CP, Jep RSA64154
Red Brome Bromus madritensis ssp. rubens

Formerly Bromus rubens

exotic EN, R, JG, M BFS, CP, Jep RSA614153

(as Bromus rubens)

UCR107774
Giant Wild-Rye Elymus condensatus

Formerly Leymus condensatus.

native SD BFS, CP, Jep
Rattail Sixweeks Grass Festuca myuros

Originally Vulpia myuros var. hirsuta.

exotic VS CP, Jep RSA612080
RSA614151
RSA706973

(all above as Vulpia myuros)

UCR108459
Goldentop Lamarckia aurea exotic S CP, Jep RSA707939
Crimson Fountaingrass Pennisetum setaceum exotic SD CP, Jep
Annual Bluegrass Poa annua exotic VS CP, Jep RSA616151
Annual Beard Grass, Rabbitfoot Grass Polypogon monspeliensis exotic VS CP, Jep RSA616145
RSA764410
Mediterranean Grass, Old Han Schismus Schismus barbatus exotic VS CP, Jep RSA616154

Pondweed Family (Potamogetonaceae)

Fennel-leaf Pondweed Stuckenia pectinata

Formerly Potamogeton pectinatus.

native EN, JG CP, Jep

Brodiaea Family (Themidaceae)

Blue Dicks Dichelostemma capitatum ssp. capitatum

Formerly placed in Liliaceae.

native JG, S BFS, CP, Jep RSA612069
RSA613091
UCR108286

Cattail Family (Typhaceae)

Broad-leaved Cattail Typha latifolia native JG CP, Jep RSA611977

CONIFERS

Pine Family (Pinaceae)

Knobcone Pine Pinus attenuata native* JG CP, Jep
Lacebark Pine Pinus bungeana exotic S
Coulter Pine Pinus coulteri native* EN, JG CP, Jep
Torrey Pine Pinus torreyana native* VS CP, Jep

FERNS

Brake Family (Pteridaceae)

Goldback Fern Pentagramma triangularis native SD BFS, CP, Jep

*Key:

Origin:
native = Native to California, according to The Jepson Manual
native* = California native planted on the BFS; not part of the indigenous flora
native† = California native naturalized on the BFS; not part of the indigenous flora
exotic = Not native to California, according to The Jepson Manual
Sources:
DC = Dan Cooper, Cooper Ecological Monitoring, Inc.
EN = Envicom, Field survey on the Robert J. Bernard Biological Field Station, September 28 and October 13, 1998
JG = Jeanne Marie Gural, Biological survey of the Robert J. Bernard Biological Field Station, Fall 1997 through Spring 2000
JW = Justin Wood, Rancho Santa Ana Botanic Garden
KH = Kristen Hasenstab, Rancho Santa Ana Botanic Garden
JS = Jane Strong, California Native Plant Society
LG = Leroy Gross, Rancho Santa Ana Botanic Garden
M = Field Station Manual, undated
R = Recon survey of the the Bernard Biological Field Station, Spring 1998
RSA = Rancho Santa Ana Botanic Garden plant list for the Robert J. Bernard Biological Field Station, Spring 1998
S = Documented field observation by Claremont Colleges faculty or staff
SD = Stephen Dreher, BFS Manager, 2001-2010
TR = Identification by Tim Ross
VS = Identification by Valerie Soza, 2001, or according to RSABG records (courtesy Valerie Soza, 2001)
Links to Photos and Descriptions:
BFS = Robert J. Bernard Biological Field Station
CP = CalPhotos, Digital Library Project, UC Berkeley. Clicking the CalPhotos link for a listing will return a display of all photos for that taxon. If you want to use any of these photos, be sure to check the guidelines for Using the Photos in CalPhotos
Jep = Jepson treatment and bioregional distribution from the Jepson eFlora, which parallels The Jepson Manual: Vascular Plants of California, 2nd Edition.
Herbarium specimens:
POM = Pomona College Herbarium Collection, now combined with the Rancho Santa Ana Botanic Garden Herbarium
RSA = Rancho Santa Ana Botanic Garden Herbarium
SEINET = Southwest Environmental Information Network
UC = The University and Jepson Herbaria, University of California, Berkeley
UCR = University of California Riverside Herbarium

© 2001-2016 Bernard Field Station Faculty Advisory Committee
Page last updated 24 October 2016 by Nancy Hamlett.